This article was republished on April 23, 2015, to correct errors in the equations that were introduced during the typesetting process. The publisher apologizes for the errors. Please download this article again to view the correct version.
All Science Journal Classification (ASJC) codes
- Ecology, Evolution, Behavior and Systematics
- Cellular and Molecular Neuroscience
- Molecular Biology
- Computational Theory and Mathematics
- Modeling and Simulation